Photosystems are the pigment-protein complexes located in the thylakoid membranes of chloroplasts. They play a riding role in light-dependent reactions of photosynthesis by capturing and then converting light energy into chemical energy. There are two types of photosystems: Photosystem 1 and Photosystem 2, which differ in their function and composition.
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Photosystems are embedded in the thylakoid membranes of chloroplasts and play a critical role in capturing the energy from light to drive electron transport, which results in the formation of ATP and NADPH. These are products essential for the Calvin cycle and glucose synthesis in plants.
The very knowledge of photosystems is the most important topic for any student from Class 9 until the undergraduate NEET students. Photosystems are a very basic concept both in plant biology and bioenergetics. The significance of photosystems is not only associated with their role in the mechanism of photosynthesis but has far-reaching applications in bioengineering or biological engineering projects like artificial photosynthesis in research, photovoltaic systems development for renewable energy, etc.
The structure of photosystems is so complex and so highly specialised that they talk about the capture and conversion of light energy to the extreme.
Each photosystem is the optimal structure for carrying out the light-involving reactions as well as absorbing the light it needs during its particular range of wavelengths.
A photosystem consists of a reaction centre chlorophyll where the primary light-induced credit to the electron transport chain originates. It is connected to an antenna complex.
Reaction center chlorophyll: P700.
Core antenna complex: 6 to 25 chlorophyll molecules and several proteins called the core antenna complex.
Primary electron acceptor: its primary electron acceptor is an iron-sulfur protein.
Location: The PSI is located in the membranes of the thylakoids.
Reaction Center Chlorophyll: P680
Core Antenna Complex: There is significant variation in the chlorophyll and carotenoid pigments.
Primary Electron Acceptor: Plastoquinone.
Oxygen-Evolving Complex: Salt in the path, which is responsible for water-splitting and evolution of oxygen.
Site: Thylakoid membrane, across the stack of grana.
Feature | Photosystem 1 (PSI) | Photosystem 2 (PSII) |
Reaction Center Chlorophyll | P700 | P680 |
Primary Function | NADPH production | Water splitting and ATP production |
Location | Stroma lamellae | Grana |
Oxygen Evolution | No | Yes |
Electron Donor | Plastocyanin | Water |
Electron Acceptor | Ferredoxin | Plastoquinone |
Light Absorption Peaks | 700 nm | 680 nm |
Role in Light-dependent Reactions
It absorbs light energy of wavelength 700 nm.
Transfers electrons to ferredoxin
Participates in generating NADPH
It absorbs light energy of wavelength 680 nm
Splits H2O molecules to release oxygen
Transfers electrons to plastoquinone
Participates in generating ATP.
The electron transport chain of photosystems is a cascade of redox steps that process light energy, eventually converting it into the chemical energy in the coenzymes ATP and NADPH.
Absorption of light by P700.
Excitation of electrons and transfer of the same to the primary acceptor.
Subsequent transport of electrons to ferredoxin.
Catalysation of the formation of NADPH by the enzyme ferredoxin-NADP+ reductase.
Absorption of light by P680.
Oxygen and protons are released.
Electrons donated to plastoquinone and passed to cytochrome b6f complex.
Photosystems may have slight variations from one type of plant to the other, making some more efficient or compatible with certain environments than others.
C3 Plants: Normal structure and functioning of PSI and PSII.
C4 Plants: More efficient photosynthesis in high-temperature environments by way of adaptations.
CAM Plants: Temporal separation of the activity of PSI and PSII to avoid water loss.
The efficiency of PSI and PSII influences the rate at which ATP and NADPH are produced.
Differences in this respect can lead to differences in growth and in the ability to adapt to environmental conditions.
Photosystem 1 produces mainly NADPH, while photosystem 2 facilitates the splitting of water and the formation of ATP.
Photosystems capture the light energy and convert it into chemical energy. This will later be in the manufacture of glucose during photosynthesis.
PSI is situated in the stroma lamellae of the thylakoid membrane. PSII, on the other hand, is located in the grana.
PSII initiates the chain that produces ATP and PSI yields electrons to NADP+ to generate NADPH.
PSI best absorbs light at 700 nm, while PSII best absorbs light at 680 nm.
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