Phloem Transport Flow From Source to Sink, Difference between Source and Sink in Plants

Phloem Transport Flow From Source to Sink, Difference between Source and Sink in Plants

Edited By Irshad Anwar | Updated on Jul 02, 2025 06:56 PM IST

Definition Of Phloem transport

It's the process of distribution of organic nutrients, mainly sugars produced in photosynthesis, in a plant. This function is attributed to phloem, a vascular tissue. Phloem transport plays a very essential role in plant physiology supplying energy and nutrients to the growing tips and roots from storage organs, hence supporting growth, development, and general health. This is the process that helps in maintaining the metabolic activities of the plant and, at the same time, enables the proper distribution of energy throughout the plant.

Mechanism Of Phloem Transport

The mechanism is described below.

Pressure Flow Hypothesis (Mass Flow Hypothesis)

Explanation of the pressure flow mechanism

The pressure flow hypothesis explains the flow of nutrients, particularly sugars like sucrose through the phloem from the source, the area where the sugars are produced, for instance, the leaf, to a sink, where they are used or stored, as in the case of roots or fruits.

Steps involved in the process

  • At the Source, Sugars are actively loaded into the sieve tube elements of the phloem, increasing the osmotic pressure.

  • Water is gained into the sieve tubes from surrounding tissues by osmosis because of the high osmotic pressure.

  • Bulk flow as turgor pressure increases, pushing the phloem sap along the sieve tubes toward the sink.

  • Repression at the Sink: Sugars are actively transported out of the sieve tubes at the sink, decreasing the osmotic pressure.

  • Water Movement: Water exits the sieve tubes at the sink. This preserves a pressure gradient that keeps the flow of the sap going.

Active And Passive Transport In Phloem

Role of active transport in loading and unloading of sugars

  • Active mechanisms of transport using energy gained from the breakdown of ATP are used in sugar uptake within the sieve tubes at the source and their removal at the sink.

  • Proton pumps, H+-ATPase, create a gradient that makes it possible for the active transport of sugars against a concentration gradient.

Role of osmosis and diffusion

  • Osmosis establishes the flow of water in and out of the phloem. This is a vital process in the maintenance of the pressure gradient responsible for the flow of sap.

  • Diffusion provides for the passive transport of water and solutes across cell membranes, thus accomplishing the overall transportation in the phloem.

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Source To Sink Transport

Source: Any vegetative body part or organ of a plant from which the sugars are produced or mobilized for transport. Such as:

  • Leaves: The photosynthetic organs where the sugars are produced.

  • Storage Organs During Mobilisation: These are such structures as tubers and bulbs when they release stored nutrients to support new growth.

Sink: Any tissue or organ that consumes or stores sugars, such as:

  • Roots: The organs themselves that need energy for growth and maintenance.

  • Fruits: The developing organs store up sugars for growth and further ripening.

  • Seeds: Nutrients are stored in the developing structures for germination.

  • Storage Organs During Accumulation: Organs like tubers and bulbs when themselves are accumulating nutrients for future use.

Seasonal Changes In Source And Sink

Spring and Early Summer: Leaves act as sources, synthesizing sugars which get translocated to the growing roots, shoots and developing fruits and sills.

Tubers and bulbs on the other hand act as sinks storing up sugars for winter.

The stored sugars in tubers and bulbs now acting as sources, mobilize for support of new growth in roots and shoots which are now acting as sinks.

Differences Between Source And Sink In Plants

Characteristic

Source

Sink

Photosynthetic Tissues

Leaves (the primary source of photosynthates)

Not typically involved in photosynthesis

Storage Organs During Mobilization

Tubers, bulbs, and other storage organs when they release stored nutrients

Not applicable

Growing Tissues

Not applicable

Roots, shoots, and developing fruits and seeds

Storage Organs During Accumulation

Not applicable

Tubers, bulbs, and other storage organs when they accumulate nutrients

Role

Produces or mobilizes sugars and nutrients

Consumes or stores sugars and nutrients

Seasonal Changes

Active during periods of growth and photosynthetic activity

Active during periods of growth, development, and nutrient storage

Examples

Leaves during photosynthesis, tubers during spring growth

Roots, fruits, seeds during development, tubers during nutrient accumulation

Function in Nutrient Cycle

Source of energy and building blocks for plant growth

Utilizes and stores energy and nutrients for growth and development


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Frequently Asked Questions (FAQs)

1. What is phloem transport in plants?

Transportation of nutrients, mainly sugars from source to sink in plants

2. What is phloem transport in plants?
Phloem transport is the process by which plants move sugars and other organic compounds from areas of production (sources) to areas of use or storage (sinks). This transport occurs through specialized vascular tissue called phloem and is essential for distributing nutrients throughout the plant.
3. What is the difference between source and sink in plants?

The source is that part of the plant, which produces or releases sugars like leaves. The sink is that part of the plant, which consumes or stores sugars like roots, fruits, or seeds.

4. How does the pressure flow hypothesis explain phloem transport?

One of the most popular ones is the pressure flow hypothesis. According to this, the flow of sugars through the phloem is due to a difference in the pressure that develops because of the active loading of the sugars at the source and their passive unloading at the sink.

5. What factors affect phloem transport in plants?

The environmental ones are those including temperature, light and availability of water; the internal ones are the plant hormones and health.

6. Why is phloem transport important for plant growth?

Phloem transport contributes crucially to the whole–plant nutrient distribution toward growth, development, and stress responses.

7. What other substances besides sugars are transported in the phloem?
While sugars (primarily sucrose) are the main substances transported in the phloem, it also carries amino acids, hormones, proteins, RNA, and various minerals. This makes phloem crucial for long-distance signaling and resource allocation within plants.
8. What is the significance of phloem transport for plant survival?
Phloem transport is essential for plant survival as it allows for the distribution of sugars and other organic compounds throughout the plant. This enables plants to support growth in non-photosynthetic tissues, respond to environmental stresses, and store energy for future use.
9. What role do companion cells play in phloem transport?
Companion cells are closely associated with sieve tube elements in the phloem and play a crucial role in phloem loading and maintenance. They provide metabolic support to the sieve tubes, assist in sugar loading, and help maintain the pressure gradient necessary for transport.
10. How does the rate of phloem transport compare to that of xylem transport?
Phloem transport is generally slower than xylem transport. While xylem can transport water at rates of several meters per hour, phloem transport typically occurs at rates of about 1 meter per hour. This difference is due to the mechanisms driving each type of transport.
11. What is a source in plant transport?
A source is any plant part that produces more sugars than it uses, typically through photosynthesis. Common sources include mature leaves, which generate excess sugars that are exported to other parts of the plant.
12. What is a sink in plant transport?
A sink is any plant part that uses more sugars than it produces. Sinks are typically growing or storage tissues that require imported sugars for growth, metabolism, or storage. Examples include roots, developing fruits, and young leaves.
13. Can a plant part be both a source and a sink?
Yes, a plant part can switch between being a source and a sink depending on its developmental stage and environmental conditions. For example, a young leaf initially acts as a sink, importing sugars for growth, but becomes a source once it matures and produces excess sugars through photosynthesis.
14. How does the direction of phloem transport differ from xylem transport?
Phloem transport is bidirectional, meaning it can move substances both up and down the plant, depending on the location of sources and sinks. In contrast, xylem transport is unidirectional, always moving water and minerals upward from roots to shoots.
15. What happens to sugars when they reach a sink?
When sugars reach a sink, they are unloaded from the phloem and either used immediately for growth and metabolism or converted into storage forms like starch. This unloading process maintains the concentration gradient that drives phloem transport.
16. What drives the flow of phloem sap from source to sink?
The flow of phloem sap is driven by a pressure gradient created by the loading of sugars at the source and unloading at the sink. This process, known as pressure flow, causes water to move into the phloem at the source and out at the sink, creating a bulk flow of sap.
17. What is the pressure flow hypothesis?
The pressure flow hypothesis, proposed by Ernst Münch in 1930, explains how phloem transport occurs. It suggests that the loading of sugars into the phloem at the source creates high osmotic pressure, drawing water in and creating hydrostatic pressure that pushes sap towards areas of lower pressure (sinks).
18. How do plants load sugars into the phloem at the source?
Plants use active transport mechanisms to load sugars into the phloem at the source. This can occur through symplastic loading (via plasmodesmata) or apoplastic loading (across cell membranes), both of which require energy in the form of ATP.
19. Why is phloem transport considered a bulk flow process?
Phloem transport is considered a bulk flow process because the entire solution (including water, sugars, and other dissolved substances) moves together in the same direction, driven by the pressure difference between source and sink.
20. How does the concentration of sugars in phloem sap compare to that in other plant cells?
The concentration of sugars in phloem sap is much higher than in other plant cells, often reaching 30% or more. This high concentration is crucial for creating the osmotic gradient that drives phloem transport.
21. How does phloem transport respond to changes in a plant's environment?
Phloem transport is highly responsive to environmental changes. For example, drought stress can alter source-sink relationships, redirecting sugars to support root growth. Similarly, changes in light availability can affect photosynthesis rates, impacting sugar production and transport patterns.
22. How do plants regulate the direction of phloem transport?
Plants regulate the direction of phloem transport by controlling the loading and unloading of sugars at sources and sinks. The strength of a sink (its demand for sugars) and the production capacity of a source determine the direction and magnitude of transport.
23. What is the role of plasmodesmata in phloem transport?
Plasmodesmata are channels that connect adjacent plant cells, allowing for direct cytoplasmic connections. In phloem transport, they play a crucial role in symplastic loading of sugars into the phloem and in the movement of substances between companion cells and sieve tube elements.
24. How does phloem transport differ between herbaceous plants and trees?
While the basic mechanism of phloem transport is similar in herbaceous plants and trees, the distance over which transport occurs is much greater in trees. This requires additional adaptations in trees, such as more efficient loading and unloading mechanisms and stronger pressure gradients.
25. What happens to phloem transport during the winter in deciduous trees?
In deciduous trees, phloem transport significantly decreases during winter as leaves fall and photosynthesis ceases. However, some transport still occurs to redistribute stored sugars from roots or stems to support essential metabolic processes and prepare for spring growth.
26. How do plants prioritize sinks when resources are limited?
When resources are limited, plants prioritize sinks based on their importance for survival and reproduction. Generally, developing fruits and seeds receive priority, followed by growing shoot tips and roots. This prioritization can change based on environmental conditions and the plant's developmental stage.
27. What is phloem unloading, and why is it important?
Phloem unloading is the process by which sugars and other substances are removed from the phloem at sink tissues. It's crucial because it maintains the concentration gradient that drives phloem transport and allows sink tissues to access the transported nutrients for growth, storage, or metabolism.
28. How does the structure of sieve tube elements facilitate phloem transport?
Sieve tube elements are specialized cells in the phloem that lack a nucleus and have porous end walls called sieve plates. This structure allows for a continuous tube through which phloem sap can flow, minimizing resistance and facilitating long-distance transport.
29. What is the relationship between photosynthesis and phloem transport?
Photosynthesis and phloem transport are closely linked. Photosynthesis produces the sugars that are transported in the phloem, while phloem transport distributes these sugars to non-photosynthetic parts of the plant. The rate of photosynthesis can directly affect the amount of sugars available for transport.
30. How do plants maintain the osmotic gradient necessary for phloem transport?
Plants maintain the osmotic gradient for phloem transport through continuous loading of sugars at the source and unloading at the sink. Additionally, companion cells actively pump protons out of the sieve tubes, which helps to maintain the electrochemical gradient that drives sugar loading.
31. What is the role of callose in phloem transport?
Callose is a polysaccharide that can rapidly form plugs in sieve plates. While it can impede phloem transport, its primary role is protective. Callose deposition occurs in response to injury or stress, sealing off damaged areas to prevent loss of phloem sap and limit the spread of pathogens.
32. How does phloem transport contribute to the carbon cycle?
Phloem transport plays a crucial role in the carbon cycle by redistributing fixed carbon (in the form of sugars) throughout the plant. This includes moving carbon to roots, where it can be released into the soil through root exudates or stored, contributing to soil carbon sequestration.
33. What is the concept of source strength in phloem transport?
Source strength refers to the capacity of a source tissue to produce and export sugars. It is determined by factors such as photosynthetic rate, leaf size, and the efficiency of phloem loading. Stronger sources can support more sinks or drive transport over longer distances.
34. How does sink strength influence phloem transport?
Sink strength is the capacity of a sink tissue to import and utilize sugars. Stronger sinks create a steeper concentration gradient, driving faster phloem transport. Factors affecting sink strength include growth rate, metabolic activity, and the efficiency of sugar unloading and utilization.
35. What is the significance of the symplast in phloem transport?
The symplast, the continuous network of cell cytoplasm connected by plasmodesmata, is crucial for phloem transport. It allows for the movement of sugars from mesophyll cells to the phloem without crossing cell membranes, facilitating efficient loading in some plants and enabling communication between cells.
36. How do viruses exploit the phloem transport system?
Many plant viruses exploit the phloem transport system to spread throughout the plant. They can enter the phloem through infected cells and then travel along with the phloem sap to infect distant parts of the plant, taking advantage of the plant's own nutrient distribution system.
37. What is the role of aquaporins in phloem transport?
Aquaporins are water channel proteins that facilitate the rapid movement of water across cell membranes. In phloem transport, they play a crucial role in maintaining the water balance necessary for pressure flow, allowing for quick adjustments in osmotic pressure at both sources and sinks.
38. How does temperature affect phloem transport?
Temperature can significantly impact phloem transport. Higher temperatures generally increase the rate of transport by enhancing metabolic activities, including sugar production and loading. However, extreme temperatures can disrupt transport by affecting membrane properties and enzyme activities.
39. What is the importance of phloem transport in crop productivity?
Phloem transport is crucial for crop productivity as it determines the distribution of sugars to economically important parts of the plant, such as fruits, grains, or tubers. Understanding and optimizing phloem transport can lead to improved crop yields and quality.
40. How do plants coordinate xylem and phloem transport?
Plants coordinate xylem and phloem transport through various mechanisms. For example, the loading of sugars into the phloem at the source creates an osmotic gradient that draws water from the xylem, linking the two systems. This coordination ensures efficient use of water and nutrients throughout the plant.
41. What is the concept of a phloem mobile signal?
A phloem mobile signal is any molecule transported in the phloem that can induce responses in distant parts of the plant. These signals can include hormones, proteins, or RNA molecules, and they play crucial roles in coordinating growth, development, and responses to environmental stimuli across the entire plant.
42. How do plants protect the sugars in phloem sap from bacterial or fungal degradation?
Plants protect phloem sap from microbial degradation through several mechanisms. These include maintaining high osmotic pressure in the phloem, which inhibits microbial growth, producing antimicrobial compounds, and using callose to seal off infected areas quickly.
43. What is the relationship between phloem transport and plant hormones?
Phloem transport and plant hormones are closely interrelated. The phloem serves as a conduit for transporting hormones throughout the plant, allowing for long-distance signaling. Conversely, hormones can influence phloem transport by affecting source and sink activities and modulating sugar loading and unloading processes.
44. How does grafting affect phloem transport?
Grafting, a common horticultural practice, can significantly impact phloem transport. When successful, the phloem of the scion (upper part) and rootstock (lower part) connect, allowing for the continuation of phloem transport across the graft union. This can lead to interesting interactions between genetically distinct tissues in terms of source-sink relationships.
45. What is phloem translocation, and how does it differ from transport?
Phloem translocation refers to the entire process of moving substances from source to sink, including loading, transport, and unloading. While phloem transport specifically describes the movement of sap through the sieve tubes, translocation encompasses the broader process of resource allocation within the plant.
46. How do plants adjust phloem transport during fruit development?
During fruit development, plants adjust phloem transport by increasing the sink strength of developing fruits. This involves hormonal changes that enhance sugar unloading and utilization in the fruit, as well as increased phloem development to support the higher demand for nutrients.
47. What is the role of sucrose synthase in phloem transport?
Sucrose synthase is an enzyme that plays a crucial role in phloem transport, particularly at sinks. It catalyzes the breakdown of sucrose into glucose and fructose, facilitating sugar unloading and utilization. This activity helps maintain the concentration gradient that drives phloem transport.
48. How does phloem transport contribute to plant defense mechanisms?
Phloem transport contributes to plant defense by facilitating the movement of defense-related compounds throughout the plant. This includes the transport of signaling molecules that can induce systemic acquired resistance, as well as the movement of defensive proteins and secondary metabolites to sites of pathogen attack.
49. What is the concept of a symplastic domain in relation to phloem transport?
A symplastic domain refers to a group of cells interconnected by plasmodesmata, allowing for direct cytoplasmic communication. In phloem transport, symplastic domains are important for coordinating loading and unloading processes, as well as for regulating the movement of substances between the phloem and surrounding tissues.
50. How do plants maintain phloem transport during drought stress?
During drought stress, plants maintain phloem transport through various adaptations. These include osmotic adjustment to maintain turgor pressure, increased loading of compatible solutes into the phloem, and prioritization of transport to essential sinks like roots to support water uptake.
51. What is the significance of phloem transport in the movement of amino acids?
Phloem transport is crucial for the movement of amino acids throughout the plant. This is particularly important for redistributing nitrogen from senescing leaves to growing tissues or storage organs. The transport of amino acids in the phloem allows plants to efficiently recycle and reallocate this essential nutrient.
52. How does the day-night cycle affect phloem transport?
The day-night cycle significantly affects phloem transport. During the day, photosynthesis increases sugar production and phloem loading in leaves, enhancing transport. At night, when photosynthesis ceases, transport rates may decrease, but movement of stored sugars from temporary storage in leaves to other parts of the plant continues.
53. What is the role of phloem transport in seed development?
Phloem transport plays a critical role in seed development by delivering sugars, amino acids, and other essential compounds to the developing seeds. This supports embryo growth, endosperm development, and the accumulation of storage compounds that will later support seedling growth.
54. How do parasitic plants exploit host plant phloem transport?
Parasitic plants exploit host plant phloem transport by forming specialized structures called haustoria that penetrate the host's vascular tissue. These structures allow the parasite to tap directly into the host's phloem, siphoning off sugars and other nutrients for their own growth and development.
55. What are the implications of understanding phloem transport for genetic engineering in agriculture?
Understanding phloem transport is crucial for genetic engineering in agriculture. It allows for the development of strategies to enhance crop yield by optimizing source-sink relationships, improving nutrient allocation to harvestable parts, and enhancing plant responses to environmental stresses. This knowledge can guide the creation of more efficient and productive crop varieties.

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