Fertilization in flowering plants is the process by which male and female gametes fuse to form a zygote.
The fertilization process is described below:
Self-pollination: This is the transfer of pollen from anther to the stigma of the same flower, or different flower of the same plant.
Cross-pollination: This is the transfer of pollen from the anther of one plant to another plant's stigma.
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A pollen grain lands on the stigma, germinates and produces a pollen tube.
The pollen tube grows down the style and into the ovary to the ovule.
One sperm combines with an egg cell to form a zygote (2n).
The other sperm combines with two polar nuclei to form the triploid endosperm (3n).
Such post-fertilization changes in a flower are necessary for the formation of seed and fruit.
This is a continuous process that finally propagates to the perpetuation of the plant life cycle.
Mitotic division of the zygote forms the embryo.
The triploid cell divides to nourish the developing embryo.
The sequence from the development of the zygote to the embryo brings about seed formation.
The stages are described below:
Initial divisions of the zygote.
The embryo becomes spherical.
The embryo becomes heart-shaped as cotyledons develop.
Additional elongation and differentiation of the embryo.
The zygote's developmental series goes through a couple of stages of development that eventually lead up to seed formation.
The development stages are given below:
Initial cell divisions of the zygote.
The embryo develops into a sphere.
The embryo takes on a heart-like shape as cotyledons develop.
Further elongation and differentiation of the embryo.
The protective outer layers develop from the integuments of the ovule.
The accumulation of nutrients.
Seeds become dehydrated and their coats harden.
The endosperm develops after fertilization and serves to nourish the embryo.
Nuclear: Free nuclear divisions followed by cell wall formation.
Cellular: Cell wall formation coincides with nuclear divisions.
Helobial: A fusion of nuclear and cellular types.
Supplies nutrition to the developing embryo.
The ovary becomes a fruit, which serves to protect and disperse the seed.
The ovary wall thickens and differentiates into pericarp layers.
Simple Fruits: Derived from a single ovary (e.g. cherry).
Aggregate Fruits: Originate from several ovaries of one flower (e.g. raspberry).
Multiple Fruits: Originate from ovaries of several flowers (e.g. pineapple).
Auxins: Stimulate cell elongation.
Gibberellins: Promote growth and development.
Cytokinins: Stimulate cell division.
Seed dispersal mechanisms contribute to the distribution of seeds for species propagation.
Wind: Seeds are light enough to become airborne (e.g. dandelion).
Water: Seeds are buoyant and distributed by water (e.g. coconut).
Animals: Seeds have either barbs or an attached nutritious reward to attract animals that disperse them (e.g. burs, berries).
Mechanical: Explosive release, for example, touch-me-not.
Special structures such as wings or flotation organs.
Spreads plants out, reduces competition, and enhances colonization of new locations.
Dormancy provides for seed survival during adverse conditions until the seed germinates.
A resting or quiescent state of metabolic inactivity.
Temperature: Low temperatures can initiate dormancy.
Moisture: Amount of available water impact.
Light: Some seeds require the action of particular light intensity to end dormancy.
Imbibition: Water uptake by the seed.
Activation of Enzymes: Stored nutrients are hydrolysed by enzymes.
Radicle and Plumule Emergence: Root and shoot development.
Correct Temperature, water, oxygen, and light.
Environment plays an essential role in determining changes after fertilisation and the process of seed development.
Action of Temperature, Water, Light, and Soil on Changes after Fertilisation
Appropriate environmental conditions are required for the development of seeds and fruits.
Effects of Biotic Factors, for example, Pests, Pollinators
Biotic interactions can affect fertilization, seed development as well as dispersal.
The post-fertilization changes are the following: from the zygote, an embryo develops; the endosperm forms and the seed and fruit mature.
The endosperm develops from the triploid cell resulting from double fertilization, providing nutrients to the developing embryo.
Seed development in flowering plants occurs through stages: proembryo stage, globular stage, heart stage, torpedo stage
The overall success of fertilization, seed development, and germination is influenced by factors such as temperature, water, light, and soil conditions.
Seed dispersal occurs through wind, water, animals, and mechanical means. All of these have some adaptations that assist in their effective dispersal.
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