Plastids are unique cell organelles present in plant cells and algae. These are well known for their crucial role in the process of photosynthesis, pigment synthesis, and storage. Plastids is a topic of the chapter Cell: The Unit of Life in Biology.
Plastids are important cell organelles in plants and algae. These carry a range of functions, such as photosynthesis, pigment synthesis, and the storage of starches, oils, and proteins. Plastids contain their DNA and the system for synthesising proteins.
Plastids are essential for normal plant cell functioning. The process of energy production and metabolism is important for developing plants, and plastids play a consequential role in it. Plastids were first observed during the 19th century when scientists found the so-called photosynthetic organelle, the chloroplast.
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There are different types of plastids, each of which performs different functions:
The green pigment chlorophyll in plants is due to the presence of chloroplasts.
These are double membrane cell organelles containing thylakoid and stroma.
Chloroplasts are significant in the photosynthesis process.
A pigment named carotenoid is present in the chloroplasts.
The colour of different fruits and vegetables like tomatoes, bell peppers, and carrots is due to the presence of chromoplasts.
These do not have the thylakoid structure like in chloroplasts.
The chromoplasts function as the significant location to store and synthesise pigments.
These do not have any pigment and are colourless organelles.
Leucoplasts have many subtypes, namely amyloplast (stores and synthesises starch), elaioplast (stores fats and oils ), and proteinoplast (stores proteins).
Their main function is to store essential substances used by the plants.
This type of plastid plays a significant role in the ageing and senescence of Leaves.
These are formed from chloroplasts during the breakdown process of photosynthesis.
The formation of this type of plastid takes place in the dark.
As the plant is exposed to light, etioplasts get converted into chromoplasts, and the process of photosynthesis starts.
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The plastid structure can be differentiated into three major components: the outer membrane, the inner membrane, and the thylakoid membrane.
The outer membrane is smooth and permeable by small molecules as well as ions, but the inner membrane is less permeable and has transport proteins regulating the passage of materials.
The inside of a chloroplast consists of interconnected thylakoid membranes that are arranged in stacks called grana, where light-dependent reactions take place during photosynthesis.
The stroma is a dense fluid inside plastids surrounding the thylakoid membranes containing enzymes and the plastid DNA and ribosomes.
Different plastids have distinct structural differences related to specialisation:
For example, chloroplasts contain a highly structured internal membrane.
Chromoplasts, specialised for pigment synthesis, have few internal membranes.
Leucoplasts contain no pigments and serve as storage organelles for starch, oils, and proteins with little internal membrane.
Gerontoplasts arise as chloroplasts age. They contain reduced thylakoid membranous structures and enzymes to recycle nutrients.
Etioplasts are those that are developed in the absence of light. Contains prolamellar bodies that are a precursor to the thylakoid membranous system.
Chloroplasts are the site of photosynthesis; they convert light energy to chemical energy, which is subsequently stored as glucose.
The photosynthetic machinery is set up in two ways: light-dependent reactions primarily occur in the thylakoid membranes.
Chlorophyll, in the presence of light, catalyses the generation of ATP and NADPH, and Calvin's cycle, which is a CO₂ fixation reaction, happens in the stroma.
The plant can then utilise the glucose both for its daily activities requiring energy and as raw material for the synthesis of other organic molecules.
Chromoplasts are the plastid type needed for synthesising and storing pigments like carotenoids and xanthophylls.
These pigments result in the conspicuous colours of fruits, flowers, and other plant organs, and all of these are necessary to attract the vectors that aid in pollination and seed dispersal.
The location of chromoplasts in petals and fruits demonstrates their function in enhancing the efficiency of pollination and aiding in the dissemination of seeds by animals attracted to colourful displays.
Leucoplasts are storage organelles mainly for starch, oils, and proteins.
They play a leading role in the growth and development of plants, acting as reservoirs for important nutrients.
The endosymbiotic theory accounts for the origin of plastids. This theory postulates that plastids arose from free-living cyanobacteria engulfed by ancestral eukaryotic cells. The genetic evidence is based on the presence of several genes, including plastid genes, indicating that plastid genomes share significant similarities with the genomes of cyanobacteria, indicating a common evolutionary ancestry.
Plastid development is marked by the differentiation of proplastids, the less developed and relatively undifferentiated plastids found in meristematic tissues. Various environmental factors, such as light, play an important role in their development into mature forms. For example, in the absence of light, proplastids can develop into etioplasts, a precursor to chloroplasts. These etioplasts finally form the chloroplasts.
The plastid DNA, also termed the plastome, is circular and compact, between 100 and 200 kb in length.
Unlike nuclear DNA, plastid DNA is less gene-dense and lacks a helical chromosome structure.
It most notably encodes proteins that mainly function in the photosynthesis process and gene expressions that include ribosomal proteins, RNA polymerases, and ATP synthase subunits.
Plastid and nuclear genomes are coordinated closely to ensure the proper functioning of plastids.
Indeed, the activity of these plastids has to be coordinated to maintain the efficiency and functioning of cellular activities.
In the realm of biotechnology, chloroplast genetic engineering is emerging as a potent tool. Scientists modify the chloroplast genome to express biopharmaceuticals like therapeutic proteins and vaccines, using the fact that chloroplasts express protein at higher levels. Since the yield is higher than in other systems and contamination by human pathogens is low, it reduces the cost and time needed for pharmaceutical production.
In agriculture, plastids are instrumental in increasing the yield of nutrients in crops. These same chloroplasts can be engineered to develop plants with an improved ability to resist many environmental stresses, like drought, salinity, and pests. More importantly, such improved crop varieties help to reduce the application of chemical pesticides and fertilisers, thus promoting sustainable agriculture.
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Plastids are the organelles present in the cells of plants and algae, responsible for photosynthesis, pigment synthesis, and nutrient storage.
Chloroplasts are the plastids responsible for photosynthesis, a feature that sets them apart from other plastids, including chromoplasts and leucoplasts, by utilising their green pigmentation, hence their capacity to convert light energy into chemical energy.
Chromoplasts of plants play an important role in synthesising and storing pigments that give colour to the fruits and flowers, hence helping in the attraction of pollinators for reproduction.
Leucoplasts play the role of starch, oils, and protein storage for the biological processes of growth and development of a plant, more particularly in storage organs like roots and seeds, enabling seed germination while storing the said energy for use by the plant.
Plastids can be genetically engineered, helping improve crops through genes responsible for high-yielding abilities, better nutritional qualities, or tolerance to biotic and abiotic stresses, which could contribute to solutions to challenges being faced in agriculture.
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